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Northern Goshawk 国家二级保护动物

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Scientific name: Accipiter gentilis

Chinese name: 苍鹰

Scientific classification

Northern Goshawk

The Northern Goshawk /ˈɡɒs.hɔːk/ (Old English: gōsheafoc, "goose-hawk"), Accipiter gentilis, is a medium-large bird of prey in the family Accipitridae, which also includes other diurnal raptors, such as eagles, buzzards and harriers.

Species: Accipiter gentilis
Genus: Accipiter
Subfamily: Accipitrinae
Family: Accipitridae
Order: Accipitriformes
English: Northern Goshawk
中文学名: 苍鹰
中文属名: 鹰属
中文亚科: 鹰亚科
中文科名: 鹰科
中文目名: 鹰形目
Protected in China: Chinese Bird Second class protected species
IUCN Red List: IUCN Red List
鸟类百科采用世界鸟类学家联合会(IOC)最新最权威的鸟类分类方法,Version 3.5
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Description

It is a widespread species that inhabits the temperate parts of the northern hemisphere. It is the only species in the Accipiter genus found in both Eurasia and North America. With the exception of Asia, it is the only species of "goshawk" in its range and it is thus often referred to (officially and unofficially, respectively) as simply the "Goshawk". It is mainly resident, but birds from colder regions migrate south for the winter. In North America, migratory goshawks are often seen migrating south along mountain ridge tops in September and October.

This species was first described by Linnaeus in his Systema naturae in 1758 under its current scientific name.

The Northern Goshawk appears on the flag of the Azores. The archipelago of the Azores, Portugal, takes its name from the Portuguese language word for goshawk, (açor), because the explorers who discovered the archipelago thought the birds of prey they saw there were goshawks; later it was found that these birds were kites or Common Buzzards (Buteo buteo rothschildi).

Description

Northern Goshawk
Juvenile (left) and adult by Louis Agassiz Fuertes
Northern Goshawk
Juvenile in flight

The Northern Goshawk is the largest member of the genus Accipiter. It is a raptor with short, broad wings and a long tail, both adaptations to manoeuvring through trees in the forests it lives and nests in. Across most of the species's range, it is blue-grey above and barred grey or white below, but Asian subspecies in particular range from nearly white overall to nearly black above. The juvenile is brown above and barred brown below. Juveniles and adults have a barred tail, with dark brown or black barring. Adults always have a white eye stripe. In North America, juveniles have pale-yellow eyes, and adults develop dark red eyes usually after their second year, although nutrition and genetics may affect eye color as well. In Europe and Asia, juveniles also have pale-yellow eyes, however adults develop orange-colored eyes.

The Northern Goshawk, like all accipiters, exhibits sexual dimorphism, where females are significantly larger than males. Males, being the smaller sex by around 10–25%, are 46–57 cm (18–22 in) long and have a 89–105 cm (35–41 in) wingspan. The female is much larger, 58–69 cm (23–27 in) long with a 108–127 cm (43–50 in) wingspan. Males average around 780 g (1.7 lb), with a range of 500 to 1,200 g (1.1 to 2.6 lb). The female can be more than twice as heavy, averaging 1,220 g (2.7 lb) with a range of 820 to 2,200 g (1.8 to 4.9 lb). Among standard measurements, the wing chord is 28.6–39 cm (11.3–15 in), the tail is 20–28 cm (7.9–11 in), the culmen is 2–2.6 cm (0.79–1.0 in) and the tarsus is 6.8–9 cm (2.7–3.5 in). In Eurasia, the species follows Bergmann's rule, specimens from the northern races generally are larger-bodied than goshawks near the southern reaches of the species range. Going on wing chord length, A. g. apache, found in Mexico to Arizona and New Mexico is the largest subspecies at an average of 36.8 cm (14.5 in) and is larger than more Northern subspecies in that continent, thus running contrary to Bergmann's rule. A. g. fujiyamae of Japan is the smallest race, at 30.9 cm (12.2 in) in wing chord length. In Europe, goshawks from Finland or of Finnish ancestry are prized as bigger than other goshawks.

The flight is a characteristic "flap flap, glide", but is sometimes seen soaring in migration, and is capable of considerable, sustained, horizontal speed in pursuit of prey with speeds of 38 mph (61 km/h) reported. Goshawks are sometimes confused with gyrfalcons especially when observed in high speed pursuit, with their wingtips drawn backward in a falcon-like profile.

In Eurasia, the male is sometimes confused with a female Sparrowhawk, but is larger, much bulkier and has relatively longer wings. In North America, juveniles are sometimes confused with the smaller Cooper's Hawk, however the juvenile goshawk displays a heavier, vertical streaking pattern on their chest and abdomen and sometimes appears to have a shorter tail due to its much larger and broader body. Although there appears to be a size overlap between small male goshawks and large female Cooper's Hawks, morphometric measurements (wing and tail length) of both species demonstrate no such overlap, although weight overlap can occur due to variation in seasonal condition and food intake at time of weighing. In North America, the Sharp-shinned Hawk is markedly smaller.

Habitat

Northern Goshawks can be found in both deciduous and coniferous forests. They seem to only thrive in areas with mature, old-growth woods and are typically found where human activity is relatively low. During nesting season, they favor tall trees with intermediate canopy coverage and small openings below for hunting. They can be found at almost any altitude, but are typically found at high elevations recently due to a paucity of extensive forests remaining in lowlands across much of its range. In winter months, the northernmost populations move down to warmer forests with lower elevations, continuing to avoid detection except while migrating. A majority of goshawks around the world remain sedentary throughout the year.

Food and hunting

This species hunts birds and mammals in a variety of woodland habitats, often utilizing a combination of speed and obstructing cover to ambush birds and mammals. Goshawks are often seen flying along adjoining habitat types, such as the edge of a forest and meadow; flying low and fast hoping to surprise unsuspecting prey. They are usually opportunistic predators, as are most birds of prey. The most important prey species are small mammals and birds found in forest habitats, in North America, this is compromised largely by grouse, American Crow, snowshoe hare, and red squirrel. Compared to many smaller Accipiter species, Northern Goshawks are less specialized as predators of birds, with up to 69% of their diet comprised by either birds or mammals depending on location. Prey species may be quite diverse, including pigeons and doves, pheasants, partridges, grouse, gulls, assorted waders, woodpeckers, corvids, waterfowl (mostly tree-nesting varieties such as the Aythya genus) and various passerines depending on the region. Mammal prey may include rabbits, hares, tree squirrels, ground squirrels, chipmunks, rats, voles, mice, weasels and shrews. Prey is often smaller than the hunting hawk, with an average prey mass of 275 g (9.7 oz) in one study of nesting birds in Minnesota. In the Netherlands, male prey averaged 277 g (9.8 oz) whereas female prey averaged 505 g (17.8 oz). However, Northern Goshawks will also occasionally kill much larger animals, up to the size of geese, raccoons, foxes and large hares, any of which can be more than twice their own weight. The Goshawk is likely a significant predator of other raptors, known prey including European Honey Buzzards, owls, smaller Accipiters and the American Kestrel.

Northern goshawks sometimes cache prey on tree branches or wedged in a crotch between branches for up to 32 hours. This is done primarily during the nestling stage.

Behavior

Northern Goshawk
Just missed him, (c.1898), G. E. Lodge.

In the spring breeding season, Northern Goshawks perform a spectacular "undulating flight display", and this is one of the best times to see this secretive forest bird. At this time, the surprisingly gull-like call of this bird is sometimes heard. Adults defend their territories fiercely from intruders, including passing humans. It is presumed that their unusually aggressive nest defense is an adaptation to tree-climbing bears species, such as the black bear in North America. Additional predators at the nest may include formidable species such as fishers, wolves, eagles and Great Horned and Eurasian Eagle Owls. Goshawks are most under threat from hatching until their fledgling stage, and are rarely threatened outside their own species once they attain adulthood. Other raptor species are attacked at nest sites, over competition for food and in territorial behavior and many are regularly displaced or even killed by the aggressive goshawk. The Northern Goshawk is considered a secretive raptor, and is rarely observed even in areas where nesting sites are common.

Breeding

Adults return to their nesting territories by March or April and begin laying eggs in April or May. Territories often encompass a variety of habitats, however the immediate nest area is often found in a mature or old-growth forest. The clutch size is usually 2 to 4, but anywhere from 1 to 5 eggs may be laid. The female is the primary incubator, although the male will sometimes take a shift to give the female a chance to eat. The male does most of the hunting for both the female and the young at the nest. The eggs average 59 mm × 45 mm (2.3 in × 1.8 in) and weigh about 60 g (2.1 oz). The incubation period can range from 28 to 38 days. The young leave the nest after about 35 days and start trying to fly another 10 days later. The young may remain in their parents' territory for up to a year of age, at which point sexual maturity is reached.

Status

In the United Kingdom and Ireland, the Northern Goshawk was extirpated in the 19th century because of specimen collectors and persecution by gamekeepers, but in recent years it has come back by immigration from Europe, escaped falconry birds, and deliberate releases. The Goshawk is now found in considerable numbers in Kielder Forest, Northumberland, which is the largest forest in Britain. The main threat to Northern Goshawks internationally today is the clearing of forest habitat on which both they and their prey depend, and in Britain illegal shooting and poisoning for game preservation when young dispersing move into farming areas.

In North America, several non-governmental conservation organizations petitioned the Department of Interior, United States Fish & Wildlife Service (1991 & 1997) to list the Goshawk as "threatened" or "endangered" under the authority of the Endangered Species Act. Both petitions argued for listing primarily on the basis of historic and ongoing nesting habitat loss, specifically the loss of old-growth and mature forest stands throughout the goshawk's known range. In both petitions, the U.S. Fish & Wildlife Service concluded that listing was not warranted, but state and federal natural resource agencies responded during the petition process with standardized and long-term goshawk inventory and monitoring efforts, especially throughout U.S. Forest Service lands in the Western U.S. The United States Forest Service (US Dept of Agriculture) has listed the goshawk as a "sensitive species", while it also benefits from various protection at the state level. In North America, the goshawk is federally protected under the Migratory Bird Treaty Act of 1918 by an amendment incorporating native birds of prey into the Act in 1972. The Northern Goshawk is also listed in Appendix II of the Convention on International Trade in Endangered Species (CITES) .

In falconry

Northern Goshawk
A falconer's bird in Scotland

The name "goshawk" is a traditional name from Anglo-Saxon gōshafoc, literally "goose hawk". The name implies prowess against larger quarry such as wild geese, but were also flown against crane species and other large waterbirds. The name "goose hawk" is somewhat of a misnomer, however, as the traditional quarry for goshawks in ancient and contemporary falconry has been rabbits, pheasants, partridge, and medium sized waterfowl. A notable exception is in records of traditional Japanese falconry, where goshawks were used more regularly on goose and crane species. In ancient European falconry literature, goshawks were often referred to as a yeoman's bird or the "cook's bird" due to their utility as a hunting partner catching edible prey, as opposed to the peregrine falcon, also a prized falconry bird, but more associated with nobleman and less adapted to a variety of hunting techniques and prey types found in wooded areas. The Northern Goshawk has remained equal to the peregrine falcon in its stature and popularity in modern falconry.

Goshawk hunting flights in falconry typically begin from the falconer's gloved hand, where the fleeing bird or rabbit is pursued in a horizontal chase. The goshawk's flight in pursuit of prey is characterized by an intense burst of speed often followed by a binding maneuver, where the goshawk, if the prey is a bird, inverts and seizes the prey from below. The goshawk, like other accipiters, shows a marked willingness to follow prey into thick vegetation, even pursuing prey on foot through brush.

References

  1. BirdLife International (2012). "Accipiter gentilis". IUCN Red List of Threatened Species. Version 2012.1. International Union for Conservation of Nature. http://www.iucnredlist.org/apps/redlist/details/106003466. Retrieved 16 July 2012.
  2. "Avibase: Eastern Goshawk". http://avis.indianbiodiversity.org/fauna-of-british-india-2nd-ed-vol-v-1928/eastern-goshawk-astur-gentilis-schvedowi.html.
  3. (Latin) Linnaeus, C (1758). Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata.. Holmiae. (Laurentii Salvii).. p. 89. "F. cera pedibusque flavis, corpore cinereo maculis fuscis cauda fasciis quatuor nigricantibus"
  4. "Northern Goshawk". Birds of Quebec. http://redpath-museum.mcgill.ca/Qbp/birds/Specpages/northerngoshawk.htm. Retrieved 2007-03-18.
  5. ^ abcdefghijkRaptors of the World by Ferguson-Lees, Christie, Franklin, Mead & Burton. Houghton Mifflin (2001), ISBN 0-618-12762-3.
  6. ^ ab Goshawk- Life History
  7. [1]
  8. [2]
  9. "Northern Goshawk". http://www.hangingrocktower.org/birds/goshawk.htm. Retrieved 2012-01-23.
  10. ^ abc B L Smithers, C W Boal, D E Andersen (2005). "Northern Goshawk diet in Minnesota: An analysis using video recording systems". Journal of Raptor Research 39 (3): 264–273. http://elibrary.unm.edu/sora/jrr/v039n03/p00264-p00273.pdf.
  11. Lewis, Stephen B.; Titus, Kimberly; Fuller, Mark R. (2006). "Northern Goshawk Diet During the Nesting Season in Southeast Alaska". Journal of Wildlife Management 70 (4): 1151. doi:10.2193/0022-541X(2006)70[1151:NGDDTN]2.0.CO;2. http://fresc.usgs.gov/products%5Cpapers%5C1547_Fuller.pdf.
  12. Accipiter gentilis – northern goshawk. Animal Diversity Web. University of Michigan
  13. Northern Goshawk (Accipter gentilis).
  14. C.Michael Hogan, ed. 2010. American Kestrel. Encyclopedia of Earth, U.S. National Council for Science and the Environment, Ed-in-chief C.Cleveland
  15. Woodbridge, B; Hargis C.D., 2006. Northern goshawk inventory and monitoring technical guide. Gen. Tech. Report WO-71. Washington, D.C.: U.S. Department of Agriculture, Forest Service.
  16. Lockwood, W B (1993). The Oxford Dictionary of British Bird Names. OUP. ISBN 978-0-19-866196-2.
  17. Jameson, E. W. Jr. 1962. The Hawking of Japan, the History and Development of Japanese Falconry. The Author, Davis. Calif. 97 p. 2
  18. ^ ab F.L. Beebe, H.M. Webster, North American Falconry and Hunting Hawks; 8th edition, 2000, ISBN 0-685-66290-X

Further reading

Identification

  • Vinicombe, Keith (2005) Getting to grips with Goshawks Birdwatch 153:29–33 (a discussion of Goshawk identification)

External links

  • Goshawk videos, photos & sounds on the Internet Bird Collection
  • Northern Goshawk Species Account – Cornell Lab of Ornithology
  • Northern Goshawk – USGS Patuxent Bird Identification InfoCenter
  • Environment Canada Goshawk page, including sound clip of Goshawk Call
  • Ageing and sexing (PDF) by Javier Blasco-Zumeta
  • "Accipiter gentilis". Integrated Taxonomic Information System. http://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=175300. Retrieved 22 February 2009.
  • The Medicine Bow National Forest (A Habitat for the Northern Goshawk)-Biodiversity Conservation Alliance

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参考资料

[1].  Northern Goshawk, from Wikipedia:  http://en.wikipedia.org/wiki/Northern_Goshawk

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